Department of Animal Ecology and Tropical Biology

Prof. Dr. Ingolf Steffan-Dewenter

Telephone: +49 (0)931 31-86947
Fax: +49 (0)931 31-84352

Room C039

University of Würzburg
Department of Animal Ecology and Tropical Biology
Biocenter - Am Hubland

97074 Würzburg, Germany

Research Interests
  • Animal ecology, population and community ecology, agroecology, tropical ecology, landscape ecology
  • Consequences of habitat fragmentation, land use intensification, climate change and invasive species on insect diversity and their biotic interactions
  • Relationship between biodiversity and ecosystem functioning across multiple spatiotemporal scales
  • Plant-pollinator interactions, crop and wild plant pollination services, honeybee ecology
  • Biological pest control, agri-environmental schemes, sustainable land use, biosafety research, landscape management
  • Conservation biology, management of protected areas

Current projects

Enhancing biodiversity-based ecosystem services to crops through optimized densities of green infrastructure in agricultural landscapes
Ingolf Steffan-DewenterEmily MartinAndrea HolzschuhElena Krimmer


LInking farmland Biodiversity to Ecosystem seRvices for effective ecological intensification
Ingolf Steffan-DewenterEmily MartinSarah Redlich

SFB 1047 

Insect timing: mechanisms, plasticity and interactions
C2 Timing of colony phenology and foraging activity in honeybees
Ingolf Steffan-DewenterStephan Härtel, Fabian Nürnberger

Assessing and monitoring impacts of genetically modified plants on agro-ecosystem
Ingolf Steffan-DewenterStephan HärtelNadja DannerKarin Steijven

DFG Research Unit FOR 1246 KilLi
Kilimanjaro Ecosystems Under Global Change
: Linking biodiversity, biotic interactions and biogeochemical ecosystem processes 
Ingolf Steffan-DewenterMarcell PetersAlice Claßen, Antonia Mayr, Henry Njovu, Jie Zhang

Status and trends of European pollinators

Ingolf Steffan-DewenterAndrea Holzschuh, Verena Riedinger, Sebastian Hopfenmüller

Conservation and management of ecosystem functions at multiple spatial scales

Ingolf Steffan-DewenterJochen KraussGudrun Schneider, Katharina Kallnik

Biodiversity Exploratories:  Arthropods II
Diversity of saproxylic arthropods and dead wood decomposition in differently managed forest types
Ingolf Steffan-DewenterEduard LinsenmairBeate Wende

Past projects

Effects of climate change on ecosystems and adaptation strategies

Ingolf Steffan-DewenterJochen KraussBernhard HoißAnnette Leingärtner

COCONUT - Understanding effects of land use changes on ecosystems to halt loss of biodiversity due to habitat destruction, fragmentation and degradation
Ingolf Steffan-DewenterJochen Krauss, Sabrina Brückmann

Effects of forest conversion and land use intensification in complex terrain on insect diversity and biological pest control

Ingolf Steffan-DewenterEmily Martin

Freisetzungsbegleitende Sicherheitsforschung an Mais mit multiplen Bt Genen zur Maiszünsler- und Maiswurzelbohrerresistenz Effekte transgener Bt-Maissorten mit multiplen Herbivorenresistenzen auf Honigbienen
Ingolf Steffan-DewenterStephan HärtelHarmen Hendriksma

Climate Change
Auswirkungen des Klimawandels auf Höhenverbreitung, trophische Interaktionen und genetische Diversität von Tagfaltern in Mittelgebirgsregionen
Ingolf Steffan-DewenterJochen Krauss

Biodiversity Exploratory
Biodiversitäts-Exploratorien - Effects of landscape context and land-use intensity on biodiversity and multitrophic interactions among plants, pollinators, herbivores, and their natural enemies
Ingolf Steffan-DewenterMichaela BellachJuliane Steckel

Curriculum Vitae

since 2010 Chair of the Department of Animal Ecology and Tropical Biology (Biocenter), University of Würzburg

2006-2010 Professor for Animal Population Ecology, University of Bayreuth

2002-2006 Assistant Professor, University of Göttingen

2002 Habilitation in Ecology

1997-2002 Research Associate, University of Göttingen

1997 Dr. rer. nat., Zoology, University of Göttingen


[in press]

Carrié R, Andrieu E, Ouin A, Steffan-Dewenter (2017) Interactive effects of landscape-wide intensity of farming practices and landscape complexity on wild bee diversity. Landscape Ecology doi:10.1007/s10980-017-0530-y. PUBLISHED ONLINE

Dainese M,  Riedinger V,  Holzschuh A, Kleijn D, Scheper J, Steffan-Dewenter I (2017) Managing trap-nesting bees as crop pollinators: spatiotemporal effects of floral resources and antagonists. Journal of Applied Ecology. DOI: 10.1111/1365-2664.12930. ONLINE FIRST

Lichtenberg EM, Kennedy CM, Kremen C, Batáry P, Berendse F, Bommarco R, Bosque-Pérez NA, Carvalheiro LG, Snyder WE, Williams NM, Winfree R, Klatt BK, Åström S, Benjamin F, Brittain C, Chaplin-Kramer R, Clough Y, Danforth B, Diekötter T, Eigenbrode SD, Ekroos J, Elle E, Freitas BM, Fukuda Y, Gaines-Day HR, Grab H, Gratton C, Holzschuh A, Isaacs R, Isaia M, Jha S, Jonason D, Jones VP, Klein A-M, Krauss J, Letourneau DK, Macfadyen S, Mallinger RE, Martin EA, Martinez E, Memmott J, Morandin L, Neame L, Otieno M, Park MG, Pfiffner L, Pocock MJO, Ponce C, Potts SG, Poveda K, Ramos M, Rosenheim JA, Rundlöf M, Sardiñas H, Saunders ME, Schon NL, Sciligo AR, Sidhu CS, Steffan-Dewenter I, Tscharntke T, Veselý M, Weisser WW, Wilson JK, Crowder DW (2017) A global synthesis of the effects of diversified farming systems on arthropod diversity within fields and across agricultural landscapes. Global Change Biology DOI: 10.1111/gcb.13714. EARLY VIEW.

Magrach A, Holzschuh A, Bartomeus I, Riedinger V, Roberts SPM, Rundlöf M, Vujić A, Wickens JB, Wickens VJ, Bommarco R, González-Varo JP, Potts SG, Smith HG, Steffan-Dewenter I, Vilà M (2017) Plant–pollinator networks in semi-natural grasslands are resistant to the loss of pollinators during blooming of mass-flowering crops. Ecography  40: DOI: 10.1111/ecog.02847. EARLY VIEW

Wende B, Gossner MM, Grass I, Arnstadt T, Hofrichter M, Floren A, Linsenmair KE, Weisser WW, Steffan-Dewenter I (2017) Trophic level, successional age and trait matching determine specialization of deadwood-based interaction networks of saproxylic beetles. Proceedings of the Royal Society B 284: 20170198. ONLINE FIRST


Gagic V, Kleijn D, Báldi A, Boros G, Bracht Jørgensen H, Elek Z, Garratt MPD, de Groot GA, Hedlund K, Kovávs-Hostyánszki A, Marini L, Martin E, Pevere I, Potts SG, Redlich S, Senapathi D, Steffan-Dewenter I, Switek S, Smith HG, Takács V, Tryjanowski P, van der Putten WH, van Gils S, Bommarco R (2017) Combined effects of agrochemicals and ecosystem services on crop yield across Europe. Ecology Letters 20: 1427–1436. doi: 10.1111/ele.12850.

Steijven K, Spaethe J, Steffan-Dewenter I, Härtel S (2017) Learning performance and brain structure of artificially-reared honey bees fed with different quantities of food. PeerJ 5:e3858. OPEN ACCESS.

Pufal G, Steffan-Dewenter I, Klein A (2017) Crop pollination services at the landscape scale. Current Opinion in Insect Science 21: 91–97.

Rosbakh S, Leingärtner A, Hoiss B, Krauss J, Steffan-Dewenter I, Poschlod P (2017) Contrasting Effects of Extreme Drought and Snowmelt Patterns on Mountain Plants along an Elevation Gradient. Frontiers in Plant Science 8: 1478. doi: 10.3389/fpls.2017.0147. OPEN ACCESS.

Danner N, Keller A, Härtel S, Steffan- Dewenter I (2017) Honey bee foraging ecology: Season but not landscape diversity shapes the amount and diversity of collected pollen. PLoS ONE 12(8): e0183716. pone.018371. OPEN ACCESS.

Dainese M, Schneider G, Krauss J, Steffan-Dewenter I (2017) Complementarity among natural enemies enhances pest suppression. Scientific Reports 7: 577-580.

Hudson LN, Newbold T, Contu S, Hill SLL, Lysenko I, De Palma A, ... , Floren A, … , Krauss J, ... , Steffan-Dewenter I, … , Purvis A (2017) The database of the PREDICTS (Projecting Responses of Ecological Diversity In Changing Terrestrial Systems) project. Ecology and Evolution 7: 145–188.

Nürnberger F, Steffan-Dewenter I, Härtel S (2017) Combined effects of waggle dance communication and landscape heterogeneity on nectar and pollen uptake in honey bee colonies. PeerJ 5: e3441. DOI 10.7717/peerj.3441. OPEN ACCESS.

Menegazzi P, Benetta ED, Beauchamp M, Schlichting M, Steffan-Dewenter I, Helfrich-Förster (2017) Adaptation of Circadian Neuronal Network to Photoperiod in High-Latitude European Drosophilids. Current Biology 27, 833–839. OPEN ACCESS.

Schellenberger Costa D, Classen A, Ferger S, Helbig-Bonitz M, Peters M, Böhning-Gaese K, Steffan-Dewenter I, Kleyer M (2017) Relationships between abiotic environment, plant functional traits, and animal body size at Mount Kilimanjaro, Tanzania. PLoS ONE 12: e0174157. OPEN ACCESS.

Classen A, Steffan-Dewenter I, Kindeketa WJ, Peters MK (2017) Integrating intraspecific variation in community ecology unifies theories on body size shifts along climatic gradients. Functional Ecology 31: 768–777.


Peters MK, Hemp A, Appelhans T,  Behler C, Classen A, Detsch F, Ensslin A, Ferger SW,  Frederiksen SB, Gebert F,  Haas M, Helbig-Bonitz M, Hemp C, Kindeketa WJ, Mwangomo E,  Ngereza C, Otte I, Roeder J,  Rutten G, Schellenberger Costa D, Tardanico J, Zancolli J, Deckert J, Eardley CD, Peters RS, Rooedel M-O,  Schleuning M, Ssymank A, Kakengi V, Zhang J, Boehning-Gaese K, Brandl R, Kalko EKV , Kleyer M, Nauss T, Tschapka M, Fischer M, Steffan-Dewenter I (2016) Predictors of elevational biodiversity gradients change from single taxa to the multi-taxa community level. Nature Communications 7: 13736. OPEN ACCESS.

Holzschuh A, Dainese M, González-Varo JP, Mudri-Stojnić S, Riedinger V, Rundlöf M, Scheper J, Wikkes JB, Wickens VJ, Bommarco R, Kleijn D, Potts SG, Roberts SPM, Smith HG, Vilà M, Vujić A, Steffan-Dewenter I (2016) Mass-flowering crops dilute pollinator abundance in agricultural landscapes across Europe. Ecology Letters 19: 1228-1236. OPEN ACCESS.

Soliveres S, van der Plas F, Manning P, Prati D, Gossner MM, Renner SC, Alt F, Arndt H, Baumgartner V, Binkenstein J, Birkhofer K, Blaser S, Blüthgen N, Boch S, Böhm S, Börschig C, Buscot F, Diekötter T, Heinze J, Hölzel N, Jung K, Klaus VH, Kleinebecker T, Klemmer S, Krauss J, Lange M, Morris EK, Müller J, Oelmann Y, Overmann J, Pašalić E, Rillig MC, Schaefer HM, Schloter M, Schmitt B, Schöning I, Schrumpf M, Sikorski J, Socher SA, Solly EF, Sonnemann I, Sorkau E, Steckel J, Steffan-Dewenter I, Stempfhuber B, Tschapka M, Türke M, Venter P, Weiner CN, Weisser WW, Werner M, Westphal C, Wilcke W, Wolters V, Wubet T, Wurst S, Fischer M, Allan E (2016) Biodiversity at multiple trophic levels is needed for ecosystem multifunctionality. Nature 536: 456-459.

Schneider G, Krauss J, Boetzl FA, Fritze MA, Steffan‑Dewenter I (2016) Spillover from adjacent crop and forest habitats shapes carabid beetle assemblages in fragmented semi‑natural grasslands. Oecologia 182: 1141-1150.

Peters MK, Peisker J, Steffan-Dewenter I, Hoiss B (2016) Morphological traits are linked to the cold performance and distribution of bees along elevational gradients. Journal of Biogeography 43: 2040–2049.

Danner N, Molitor AM, Schiele S, Härtel S, Steffan‐Dewenter I (2016) Season and landscape composition affect pollen foraging distances and habitat use of honey bees. Ecological Applications 26: 1920–1929.

Beer K, Steffan‑Dewenter I, Härtel S, Helfrich‑Förster C (2016) A new device for monitoring individual activity rhythms of honey bees reveals critical effects of the social environment on behavior. Journal of Comparative Physiology A 202: 555–565. OPEN ACCESS.

De Palma A, (...), Steffan-Dewenter I et al (2016) Predicting bee community responses to land-use changes: Effects of geographic and taxonomic biases. Scientific Reports 6, 31153. DOI: 10.1038/srep31153. OPEN ACCESS.

Gossner MM, Wende B, Levick S, Schall P, Floren A, Linsenmair KE, Steffan-Dewenter I, Schulze E-D, Weisser WW (2016) Deadwood enrichment in European forests – Which tree species should be used to promote saproxylic beetle diversity? Biological Conservation 201: 92–102.

Stangler ES, Hanson PE, Steffan-Dewenter I (2016) Vertical diversity patterns and biotic interactions of trap-nesting bees along a fragmentation gradient of small secondary rainforest remnants. Apidologie 47: 527–538.

Martin EA, Seo B, Park C-R, Reineking B, Steffan-Dewenter I (2015) Scale-dependent effects of landscape composition and configuration on natural enemy diversity, crop herbivory and yields. Ecological Applications 26: 448-462.

Steijven K, Steffan-Dewenter I, Härtel S (2016) Testing dose-dependent effects of stacked Bt maize pollen on in vitro-reared honey bee larvae. Apidologie 47: 216–226.


Garibaldi LA, Bartomeus I, Bommarco R, Klein AM, Cunningham SA, Aizen MA Boreux V, Garratt MPD, Carvalheiro LG, Kremen C, Morales CL, Schüepp C, Chacoff NP, Freitas BN, Gagic V, Holzschuh A, Klatt BK, Krewenka KM, Krishnan S, Mayfield MM, Motzke I, Otieno M, Petersen J, Potts SG, Ricketts TH, Rundlöf M, Sciligo A, Allesh Sinu P, Steffan-Dewenter I, Taki H, Tscharntke T, Vergara CH, Viana BF, Woyciechowski M (2015) Trait matching of flower visitors and crops predicts fruit set better than trait diversity. Journal of Applied Ecology 52: 1436–1444.

Riedinger V, Mitesser O, Hovestadt T, Steffan-Dewenter I, Holzschuh A (2015): Annual dynamics of wild bee densities: attractiveness and productivity effects of oilseed rape. Ecology 96: 1351-1360.

Keller A,  Sickel W, Ankenbrand M, Steffan-Dewenter I (2015) NGS pollen metabarcoding and automatic taxonomic classification. Genome 58: 237.

Classen A, Peters MK, Kindeketa WJ, Appelhans T, Eardley CD, Gikungu MW, Hemp A, Nauss T, Steffan-Dewenter I (2015) Temperature versus resource constraints: which factors determine bee diversity on Mount Kilimanjaro, Tanzania? Global Ecology and Biogeography 24: 642-652.

Müller J, Wende B, Strobl C, Eugster M, Gallenberger I, Floren A, Steffan-Dewenter I, Linsenmair KE, Weisser WW, Gossner MM (2015) Forest management and regional tree composition drive the host preference of saproxylic beetle communities. Journal of Applied Ecology 52: 753-762.

Kuppler J, Fricke J, Hemp C, Steffan-Dewenter I, Peters MK (2015) Conversion of savannah habitats to small-scale agriculture affects grasshopper communities at Mt. Kilimanjaro, Tanzania. Journal of Insect Conservation 19: 509-518.

Sickel W, Ankenbrand MJ, Grimmer G, Holzschuh A, Härtel S, Lanzen J, Steffan‑Dewenter I, Keller A (2015) Increased efficiency in identifying mixed pollen samples by meta‑barcoding with a dual‑indexing approach. BMC Ecology 15: 20-28. OPEN ACCESS.

Scheper J, Bommarco R, Holzschuh A, Potts SG, Riedinger V, Roberts SPM, Rundlöf M, Smith HG, Steffan‐Dewenter I, Wickens JB, Wickens VJ, Kleijn D (2015) Local and landscape‐level floral resources explain effects of wildflower strips on wild bees across four European countries. Journal of Applied Ecology 52: 1165-1175.

Gámez-Virués S, Perovic DJ, Gossner MM, Börschig C, Blüthgen N, de Jong H, Simons NK, Klein A-M, Krauss J, Maier G, Scherber C, Steckel J, Rothenwöhrer C, Steffan-Dewenter I, Weiner CN, Weisser W, Werner M, Tscharntke T, Westphal C (2015) Landscape simplification filters species traits and drives biotic homogenization. Nature Communications 6:8568. DOI: 10.1038/ncomms9568. OPEN ACCESS.

Stevenson TJ, Visser ME, Arnold W, Barrett P, Biello S, Dawson A, Denlinger DL, Dominoni D, Ebling FJ, Elton S, Evans N, Ferguson HM, Foster RG, Hau M, Haydon DT, Hazlerigg DG, Heideman P, Hopcraft JGC, Jonsson NN, Kronfeld-Schor N, Kumar V, Lincoln GA, MacLeod R, Martin SAM, Martinez-Bakker M, Nelson RJ, Reed T, Robinson JE, Rock D, Schwartz WJ, Steffan-Dewenter I, Tauber E, Thackeray SJ, Umstatter C, Yoshimura T, Helm B (2015) Disrupted seasonal biology impacts health, food security and ecosystems. Proceedings of the Royal Society B 282: 20151453.

Hoiss B, Krauss J, Steffan-Dewenter I (2015) Interactive effects of elevation, species richness and extreme climatic events on plant–pollinator networks. Global Change Biology 21: 4086–4097.

Schneider G, Krauss J, Riedinger V, Holzschuh A, Steffan-Dewenter I (2015) Biological pest control and yields depend on spatial and temporal crop cover dynamics. Journal of Applied Ecology 52: 1283–1292.

Konopik O, Steffan-Dewenter I, Grafe TU (2015) Effects of Logging and Oil Palm Expansion on Stream Frog Communities on Borneo, Southeast Asia. Biotropica 47: 636-643.

Martin EA, Reineking B, Bumsuk S, Steffan-Dewenter (2015) Pest control of aphids depends on landscape complexity and natural enemy interactions. PeerJ 3:e1095; DOI 10.7717/peerj.1095. OPEN ACCESS.

Kleijn D, Winfree R, Bartomeus I, Carvalheiro LG, Henry M, Isaacs R,  Klein A-M, Kremen C, M’Gonigle LK, Rader R, Ricketts TH, Williams NM, Adamson NL, Ascher JS, Báldi A, Batáry P, Benjamin F, Biesmeijer JC, Blitzer EJ, Bommarco R, Brand MR, Bretagnolle V, Button L, Cariveau DP, Chifflet R, Colville JF, Danforth BN, Elle E, Garratt MPD, Herzog F, Holzschuh A, Howlett BG, Jauker F, Jha S, Knop E, Krewenka KM, Le Féon V, Mandelik Y, May EA, Park MG, Pisanty G, Reemer M, Riedinger V, Rollin O, Rundlöf M, Sardinas HS, Scheper J, Sciligo AR, Smith HG, Steffan-Dewenter I, Thorp R, Tscharntke T, Verhulst J, Viana BF, Vaissière BE, Veldtman R, Westphal C, Potts SG (2015) Delivery of crop pollination services is an insufficient argument for wild pollinator conservation. Nature Communications 36: 7414.

Habel JC, Brückmann SV, Krauss J, Schwarzer J, Weig A, Husemann M, Steffan-Dewenter I (2015) Fragmentation genetics of the grassland butterfly Polyommatus coridon: stable genetic diversity or extinction debt? Conservation Genetics 16: 549-558.

Gagic V, Bartomeus I, Jonsson T, Taylor A, Winqvist C, Fischer C, Slade EM, Steffan-Dewenter I, Emmerson M, Potts SG, Tscharntke T, Weisser W, Bommarco R (2015) Functional identity and diversity of animals predict ecosystem functioning better than species-based indices. Proceedings of the Royal Society B 282: 20142620.

Stangler ES, Hanson PE, Steffan-Dewenter I (2015) Interactive effects of habitat fragmentation and microclimate on trap-nesting Hymenoptera and their trophic interactions in small secondary rainforest remnants. Biodiversity and Conservation 24: 563–577.

Keller A , Danner N, Grimmer G, Ankenbrand M, von der Ohe K, von der Ohe W, Rost S, Härtel S, Steffan-Dewenter I (2015) Evaluating multiplexed next-generation sequencing as a method in palynology for mixed pollen samples. Plant Biology 17: 558–566.


Peters MK, Mayr A, Röder J, Sanders NJ, Steffan-Dewenter I (2014) Variation in nutrient use in ant assemblages along an extensive elevational gradient on Mt Kilimanjaro. Journal of Biogeography 41: 2245–2255.

Danner N, Härtel S, Steffan-Dewenter I (2014) Maize pollen foraging by honey bees in relation to crop area and landscape context. Basic and Applied Ecology 15: 677-684.

Konopik O, Gray CL, Grafe TU, Steffan-Dewenter I, Fayle TM (2014) From rainforest to oil palm plantations: Shifts in predator population and prey communities, but resistant interactions. Global Ecology and Conservation 2: 385–394. OPEN ACCESS

Zancolli G, Rödel M-O, Steffan-Dewenter I, Storfer A (2014) Comparative landscape genetics of two river frog species occurring at different elevations on Mount Kilimanjaro. Molecular Ecology23: 4989-5002.

Hopfenmüller S, Steffan-Dewenter I, Holzschuh A (2014) Trait-Specific Responses of Wild Bee Communities to Landscape Composition, Configuration and Local Factors. PLoS ONE 9: e104439. OPEN ACCESS.

Clough Y, Ekroos J, Báldi A, Batáry P, Bommarco R, Gross N, Holzschuh A, Hopfenmüller S, Knop E, Kuussaari M, Lindborg R, Marini L, Öckinger E, Potts SG, Pöyry J, Roberts SPM, Steffan-Dewenter I, Smith HG (2014) Density of insect-pollinated grassland plants decreases with increasing surrounding land-use intensity. Ecology Letters 17: 1168–1177.

Steckel J, Westphal C, Peters MK, Bellach M, Rothenwoehrer C, Erasmi S, Scherber C, Tscharntke T, Steffan-Dewenter I (2014) Landscape composition and configuration differently affect trap-nesting bees, wasps and their antagonists. Biological Conservation 172: 56–64.

Härtel S, Steffan-Dewenter I (2014) Ecology: Honey Bee Foraging in Human-Modified Landscapes. Current Biology 24: R524-R526.

Bartomeus I, Potts SG, Steffan-Dewenter I, Vaissiere BE, Woyciechowski M, Krewenka KM, Tscheulin T, Roberts SPM, Szentgyörgyi H, Westphal C, Bommarco R (2014) Contribution of insect pollinators to crop yield and quality varies with agricultural intensification. PeerJ2: e328. OPEN ACCESS.

Leingärtner A, Hoiss B, Krauss J, Steffan-Dewenter I (2014) Combined Effects of Extreme Climatic Events and Elevation on Nutritional Quality and Herbivory of Alpine Plants. PLoS ONE 9: e93881. OPEN ACCESS.

Riedinger V, Renner M, Rundlöf M, Steffan-Dewenter I, Holzschuh A (2014) Early mass-flowering crops mitigate pollinator dilution in late-flowering crops. Landscape Ecology 29: 425-435.

Breeze TD, Vaissière BE, Bommarco R, Petanidou T, Seraphides N, Kozák L, Scheper J, Biesmeijer JC, Kleijn D, Gyldenkærne S, Moretti M, Holzschuh A, Steffan-Dewenter I, Stout JC, Pärtel M, Zobel M, Potts SG (2013) Agricultural policies exacerbate pollination service supply-demand mismatches across Europe. PLoS One 9: e82996. OPEN ACCESS.

Zancolli G, Steffan-Dewenter I, Rödel M-O (2014) Amphibian diversity on the roof of Africa: unveiling the effects of habitat degradation, altitude and biogeography. Diversity and Distributions 20: 297-308.

Marini L, Öckinger E, Bergman K-O, Jauker B, Krauss J, Kuussaari M, Pöyry J, Smith HG, Steffan-Dewenter I, Bommarco R (2014) Contrasting effects of habitat area and connectivity on evenness of pollinator communities. Ecography 37: 544-551.

Classen A, Peters MK, Ferger SW, Helbig-Bonitz M, Schmack JM, Maassen G, Schleuning M, Kalko EKV, Böhning-Gaese K, Steffan-Dewenter I (2014) Complementary ecosystem services provided by pest predators and pollinators increase quantity and quality of coffee yields. Proceedings of the Royal Society B 281: 20133148.

Leingärtner A, Krauss J, Steffan‑Dewenter I (2014): Species richness and trait composition of butterfly assemblages change along an altitudinal gradient. Oecologia 175: 613–623.

Leingärtner A, Krauss J, Steffan-Dewenter I (2014) Elevation and experimental snowmelt manipulation affect emergence phenology and abundance of soil-hibernating arthropods. Ecological Entomology 39: 412-418.

Allan E, Bossdorf O, Dormann CF, Prati D, Gossner MM, Tscharntke T, Blüthgen N, Bellach M, Birkhofer K, Boch S, Böhm S, Börschig C, Chatzinotas, Christ S, Daniel R, Diekötter T, Fischer C, Friedl T, Glaser K, Hallmann C, Hodac L, Hölzel N, Jung K, Klein A-M, Klaus, VH, Kleinebecker T, Krauss J, Lange M, Morris EK, Müller J, Nacke H, Pasalic E, Rillig MC, Rothenwöhrer C, Schall P, Scherber C, Schulze W, Socher SA, Steckel J, Steffan-Dewenter I, Türke M, Weiner CN, Werner M, Westphal C, Wolters V, Wube T, Gockel S, Gorke M, Hemp A, Renner SC, Schöning I, Pfeiffer S, König-Ries B, Buscot F, Linsenmair KE, Schulze E-D, Weisser WW, Fischer M (2014) Interannual variation in land-use intensity enhances grassland multidiversity. PNAS 111: 308-313.


Kang W, Hoffmeister M, Martin E, Steffan-Dewenter I, Han D, Lee D (2013) Effects of management and structural connectivity on the plant communities of organic vegetable field margins in South Korea. Ecological Research 28: 991-1002.

González-Varo JP, Biesmeijer JC, Bommarco R, Potts SG, Schweiger O, Smith HG, Steffan-Dewenter I, Szentgyörgyi H, Woyciechowski M, Vila M (2013) Combined effects of global change pressures on animal-mediated pollination. Trend in Ecology & Evolution 28: 524-530.

Keller A, Grimmer, Steffan-Dewenter I (2013) Diverse Microbiota Identified in Whole Intact Nest Chambers of the Red Mason Bee Osmia bicornis (Linnaeus 1758). PLoS ONE 8: e78296. OPEN ACCESS

Wagner K, Krauss J, Steffan-Dewenter I (2013): Butterfly diversity and historical land cover change along an altitudinal gradient. Journal of Insect Conservation 17: 1039-1046.

Schneider G, Krauss J, Steffan-Dewenter (2013): Predation rates on semi-natural grasslands depend on adjacent habitat type. Basic and Applied Ecology 14: 614–621.

Kennedy CM, Lonsdorf E, Neel MC, Williams NM, Ricketts TH, Winfree R, Bommarco R, Brittain C, Burley AL, Cariveau D, Carvalheiro LG, Chacoff NP, Cunningham  SA, Danforth BN, Dudenhöffer J-H, Elle E, Gaines HR, Garibaldi LA, Gratton C, Holzschuh A, Isaacs R, Javorek SK, Jha S, Klein AM, Krewenka K, Mandelik Y, Mayfield MM, Morandin L, Neame LA, Otien M, Park M, Potts SG, Rundlöf M, Saez A, Steffan-Dewenter I, Taki H, Viana  BF, Westphal C, Wilson JK, Greenleaf SS, Kremen C (2013) A global quantitative synthesis of local and landscape effects on wild bee pollinators in agroecosystems. Ecology Letters 16: 584-599.

Garibaldi LA, Steffan-Dewenter I, Winfree R, Aizen MA, Bommarco R, Cunningham SA, Kremen, C, Carvalheiro LG, Harder LD, Afik O, Bartomeus I, Benjamin F, Boreux V, Cariveau D, Chacoff NP, Dudenhöffer JH, Freitas BM, Ghazoul J, Greenleaf S, Hipólito J, Holzschuh A, Howlett B, Isaacs R, Javorek SK, Kennedy CM, Krewenka K, Krishnan S, Mandelik Y, Mayfield MM, Motzke I, Munyuli T, Nault BA, Otieno M, Petersen J, Pisanty G, Potts SG, Rader R, Ricketts TH, Rundlöf M, Seymour CL, Schüepp C, Szentgyörgyi H, Taki H, Tscharntke T, Vergara CH, Viana BF, Wanger TC, Westphal C, Williams N, Klein AM (2013) Wild Pollinators Enhance Fruit Set of Crops Regardless of Honey Bee Abundance. Science 339: 1608-1611. 

Holzschuh A, Dormann CF, Tscharntke T, Steffan-Dewenter I (2013) Mass-flowering crops enhance wild bee abundance. Oecologia172: 477-484.

Hendriksma HP, Küting M, Härtel S, Näther A, Dohrmann AB, Steffan-Dewenter I, Tebbe CC (2013) Effect of stacked insecticidal cry proteins from maize pollen on nurse bees (Apis mellifera carnica) and their gut bacteria. Plos One 8: e59589. OPEN ACCESS

Cornelius C, Leingärtner A, Hoiss B, Krauss J, Steffan-Dewenter I, Menzel A (2013) Phenological response of grassland species to manipulative snowmelt and drought along an altitudinal gradient. Journal of Experimental Botany 64: 241-251. OPEN ACCESS

Jauker B, Krauss J, Jauker F, Steffan-Dewenter I (2013) Liking life history traits to pollinator loss in fragmented calcareous grasslands. Landscape Ecology 28: 107–120.

Hoiss B, Gaviria J, Leingärtner A, Krauss J, Steffan-Dewenter I (2013) Combined effects of climate and management on plant diversity and pollination type in alpine grasslands. Diversity and Distributions 19: 386-395.

Martin EA, Reineking B, Seo B, Steffan-Dewenter I (2013) Natural enemy interactions constrain pest control in complex agricultural landscapes. Proceedings of the National Academy of Sciences of the United States of America 110: 5534–5539.


Hendriskma HP, Haertel S, Babendreier D, von der Ohe W, Steffan-Dewenter I (2012) Effects of multiple Bt proteins and GNA lectin on in vitro-reared honey bee larvae. Apidologie 43: 549-560.

Kessler M, Hertel D, Jungkunst HF, Kluge J, Abrahamczyk S, Bos M, Buchori D, Gerold G, Gradstein SR, Köhler S, Leuschner C, Moser G, Pitopang R, Saleh S, Schulze CH, Sporn SG, Steffan-Dewenter I, Tjitrosoedirdjo SS, Tscharntke T (2012) Can Joint Carbon and Biodiversity Management in Tropical Agroforestry Landscapes Be Optimized? PLoS ONE 7: e47192. OPEN ACCESS

Nielsen A, Dauber J, Kunin WE, Lamborn E, Jauker B, Moora M, Potts SG, Reitan T, Roberts S, Sõber V, Settele J, Steffan-Dewenter I, Stout JC, Tscheulin T, Vaitis M, Vivarelli D, Biesmeijer JC, Petanidou T (2012) Pollinator community responses to the spatial population structure of wild plants: A pan-European approach. Basic and Applied Ecology 13: 489-499. 

Hoiss B, Krauss J, Potts SG, Roberts S, Steffan-Dewenter I (2012) Altitude acts as an enviromental filter on phylogenetic composition, traits and diversity in bee communities. Proceedings of The Royal Society B 279: 4447-4456.

Jauker F, Bondarenko B, Becker HC, Steffan-Dewenter I (2012) Pollination efficiency of wild bees and hoverflies provided to oilseed rape. Agricultural and Forest Entomology 14: 81-87.

Tscharntke T, Tylianakis J, Rand T, Didham R, Fahrig L, Batary P, Bengtsson J, Clough Y, Crist T, Dormann C, Ewers R, Holt R, Holzschuh A, Klein A-M, Kremen C, Landis D, Laurance W, Lindenmayer D, Scherber C, Sodhi N, Steffan-Dewenter I, Thies C, van der Putten W, Westphal C (2012) Landscape moderation of biodiversity patterns and processes - eight hypotheses. Biological Reviews 87: 661-685.

Öckinger E, Bergman K-O, Franzén M, Kadlec T, Krauss J, Kuussaari M, Pöyry J, Smith HG, Steffan-Dewenter I, Bommarco R (2012) The landscape matrix modifies the effect of habitat fragmentation in grassland butterflies. Landscape Ecology 27: 121–131.

Breitbach N, Tillmann S, Schleuning M, Grünewald C, Laube I, Steffan-Dewenter I, Böhning-Gaese K (2012) Influence of habitat complexity and landscape configuration on pollination and seed-dispersal interactions of wild cherry trees. Oecologia 168: 425-437.


Hendriskma HP, Haertel S, Steffan-Dewenter I (2011) Testing Pollen of Single and Stacked Insect-Resistant Bt-Maize on In vitro Reared Honey Bee Larvae. PLoS ONE 6: e28174. OPEN ACCESS

Nielsen A, Steffan-Dewenter I, Westphal C, Messinger O, Potts S, Roberts S, Settele J, Szentgyörgyi H, Vaissière B, Vaitis M, Woyciechowski M, Bazos I, Biesmeijer J, Bommarco R, Kunin W, Tscheulin T, Lamborn E, Petanidou T (2011) Assessing bee species richness in two Mediterranean communities: importance of habitat type and sampling techniques. Ecological Research 26: 969-983.

Clough Y, Barkmann J, Juhrbandt J, Kessler M, Wanger TC, Anshary A, Buchori D, Cicuzza D, Darras K, Putra DD, Erasmi S, Pitopang R, Schmidt C, Schulze CH, Seidel D, Steffan-Dewenter I, Stenchly K, Vidal S, Weist M, Wielgoss AC, Tscharntke T (2011) Combining high biodiversity with high yields in tropical agroforests. Proceedings of the National Academy of Sciences of the United States of America 108: 8311-8316.

Brückmann SV, Krauss J, Steffan-Dewenter I (2011) The impact of habitat fragmentation on trophic interactions of the monophagous butterfly Polyommatus coridonJournal of Insect Conservation 15: 707-714. 

Garibaldi LA, Steffan-Dewenter I, Kremen C, Morales JM, Bommarco R, Cunningham SA, Carvalheiro LG, Chacoff NP, Dudenhöffer JH, Greenleaf SS, Holzschuh A, Isaacs R, Krewenka K, Mandelik Y, Mayfield MM, Morandin LA, Potts SG, Ricketts TH, Szentgyörgyi H, Viana BF, Westphal C, Winfree R, Klein A-M (2011) Stability of pollination services decreases with isolation from natural areas despite honey bee visits. Ecology Letters 14: 1062-1072.

Hendriskma HP, Haertel S, Steffan-Dewenter I (2011) Honey bee risk assessment: New approaches for in vitro larvae rearing and data analyses. Methods in Ecology and Evolution 2: 509-517.

Holzschuh A, Dormann CF, Tscharntke T, Steffan-Dewenter I (2011) Expansion of mass-flowering crops leads to transient pollinator dilution and reduced wild plant pollination. Proceedings of the Royal Society B 278: 3444-3451. OPEN ACCESS

Kessler M, Abrahamczyk S, Bos M, Buchori D, Dwi Putra D, Gradstein RS, Höhn P, Kluge J, Orend F, Pitopang R, Saleh S, Schulze CH, Sporn SG, Steffan-Dewenter I, Tjitrosoedirdjo SS, Tscharntke T (2011) Cost-effectiveness of plant and animal biodiversity indicators in tropical forest and agroforest habitats. Journal of Applied Ecology 48: 330–339.

Krauss J, Gallenberger I, Steffan-Dewenter I (2011) Decreased Functional Diversity and Biological Pest Control in Conventional Compared to Organic Crop Fields. PLoS ONE 6: e19502. OPEN ACCESS

Potts SG, Biesmeijer JC, Bommarco R, Felicioli A, Fischer M, Jokinen P, Kleijn D, Klein A-M, Kunin WE, Neumann P, Penev LD, Petanidou T, Rasmont P, Roberts SPM, Smith HG, Sørensen PB, Steffan-Dewenter I, Vaissière BE, Vilà M, Vujic A, Woyciechowski M, Zobe M, Settele J, Schweiger S (2011) Developing European conservation and mitigation tools for pollination services: approaches of the STEP (Status and Trends of European Pollinators) project. Journal of Apicultural Research 50: 152-164.

Wagner KD, Krauss J, Steffan-Dewenter, I (2011) Changes in the life history traits of the European Map butterfly Araschnis levana (Lepidoptera: Nymaphalidae) with increase in altitude. European Journal of Entomology 108: 447-452.


Brückmann SV, Krauss J, Steffan-Dewenter I (2010) Butterfly and plant specialists suffer from reduced connectivity in fragmented landscapes. Journal of Applied Ecology 47: 799-809.

Bartomeus I, Vilà M, Steffan-Dewenter I (2010) Combined effects of Impatiens glandulifera invasion and landscape structure on native plant pollination. Journal of Ecology 98: 440-450.

Bommarco R, Biesmeijer JC, Meyer B, Potts SG, Pöyry J, Roberts SPM, Steffan-Dewenter I, Öckinger E (2010) Dispersal capacity and diet breadth modify the response of wild bees to habitat loss. Proceedings of the Royal Society of London B 277: 2075–2082.

Breitbach N, Laube I, Steffan-Dewenter I, Böhning-Gaese K (2010) Bird diversity and seed dispersal along a human land-use gradient: high seed removal in structurally simple farmland. Oecologia 162: 965-976.

Dauber J, Biesmeijer JC, Gabriel D, Kunin WE, Lamborn E, Meyer B, Nielsen A, Potts SG, Roberts SPM, Sõber V, Settele J, Steffan-Dewenter I, Stout JC, Teder T, Tscheulin T, Vivarelli D, Petanidou T (2010) Effects of patch size and density on flower visitation and seed set of wild plants: a pan-European approach. Journal of Ecology 98: 188-196.

Hammen V Biesmeijer JC, Bommarco R, Budrys E, Christensen TR, Fronzek S, Grabaum R, Jaksic P, Klotz S, Kramarz P, Kröel-Dulay G, Kühn I, Mirtl M, Moora M, Petanidou T, Pino J, Potts SG, Rortais A, Schulze CH, Steffan-Dewenter I, Stout JC, Szentgyörgyi H, Vighi M, Vujic A, Westphal C, Wolf T, Zavala G, Zobel M, Settele J, Kunin WE (2010) Establishment of a cross-European field site network in the ALARM project for assessing large-scale changes in biodiversity. Environmental Monitoring & Assessment 164: 337-348.

Hoehn P, Steffan-Dewenter I, Tscharntke T (2010) Relative contribution of agroforestry, rainforest and openland to local and regional bee diversity. Biodiversity and Conservation 19: 2189-2200.

Holzschuh A, Steffan-Dewenter I, Tscharntke T (2010) How do landscape composition and configuration, organic farming and fallow strips affect the diversity of bees, wasps and their parasitoids? Journal of Animal Ecology 79: 491-500.

Henle K, Kunin W, Schweiger O, Schmeller DS, Grobelnik V, Matsinos Y, Pantis J, Penev L, Potts SG, Ring I, Similä J, Tzanopoulos J, van den Hove S, Baguette M, Clobert J, Excoffier L, Framstad E, Grodzinska-Jurczak M, Lengyel S, Marty P, Moilanen A, Porcher E, Storch D, Steffan-Dewenter I, Sykes MT, Zobel M, Settele J (2010) Securing the Conservation of Biodiversity across Adminis trative Levels and Spatial, Temporal, and Ecological Scales – Research Needs and Approaches of the SCALES Project, GAIA 19: 187-193.

Krauss J, Bommarco R, Guardiola M, Heikkinen RK, Helm A, Kuussaari M, Lindborg R, Öckinger E, Pärtel M, Pino J, Pöyry J, Raatikainen KM, Sang A, Stefanescu C, Teder T, Zobel M, Steffan-Dewenter I (2010): Habitat fragmentation causes immediate and time-delayed biodiversity loss at different trophic levels. Ecology Letters 13: 597-605.


Carré G, Roche P, Chifflet R, Morison N, Bommarco R, Harrison-Cripps J, Krewenka K Potts SG, Roberts SPM, Rodet G, Settele J, Steffan-Dewenter I, Szentgyörgyi H, Tscheulin T, Westphal C, Woyciechowski M, Vaissière BE (2009) Landscape context and habitat type as drivers of bee diversity in European annual crops. Agriculture, Ecosystems & Environment 133: 40-47.

Höhn P, Steffan-Dewenter I, Buchori D, Tscharnkte T (2009) Spatiotemporal density patterns of the pest predator Rhynchium haemorrhoidale (F.) along a land-use gradient in cacao agroforestry systems. Agroforestry Systems 76: 163-171.

Holzschuh A, Steffan-Dewenter I, Tscharntke T (2009) Grass strip corridors in agricultural landscapes enhance nest-site colonization by solitary wasps. Ecological Applications 19: 123-132.

Kessler M, Abrahamczyk S, Bos MM, Buchori D, Putra DD, Gradstein SR, Höhn P, Kluge J, Orend F, Pitopang R, Saleh S, Schulze CH, Sporn SG, Steffan-Dewenter I, Tjitrosoedirdjo SS, Tscharntke T (2009) Alpha and beta diversity of plants and animals along a tropical land-use gradient. Ecological Applications 19: 2142-2156.

Krauss J, Alfert T, Steffan-Dewenter I (2009) Habitat area but not habitat age determines wild bee richness in limestone quarries. Journal of Applied Ecology 46: 194–202.

Kuussaari M, Bommarco R, Heikkinen RK, Helm A, Krauss J, Lindborg R, Öckinger E, Pärtel M, Pino J, Rodà F, Stefanescu C, Teder T, Zobel M, Steffan-Dewenter I (2009) Extinction debt: a challenge for biodiversity conservation. TRENDS in Ecology and Evolution 24: 564-571.

Meyer B, Jauker F, Steffan-Dewenter I (2009) Contrasting resource-dependent responses of hoverfly richness and density to landscape structure. Basic and Applied Ecology 10: 178-186.

Sobek S, Goßner MM, Scherber C, Steffan-Dewenter I, Tscharntke T (2009) Tree diversity drives abundance and spatiotemporal beta-diversity of true bugs (Heteroptera). Ecological Entomology 34: 772-782.

Sobek S, Scherber C, Steffan-Dewenter I, Tscharntke T (2009) Sapling herbivory, invertebrate herbivores and predators across a natural tree diversity gradient in Germany’s largest connected deciduous forest. Oecologia 160: 279-288.

Sobek S, Steffan-Dewenter I, Scherber C, Tscharntke T (2009) Spatiotemporal changes of beetle communities across a tree diversity gradient. Diversity and Distributions 15: 660-670.

Sobek S, Tscharntke T, Scherber C, Schiele S, Steffan-Dewenter I (2009) Canopy vs. understory: Does tree diversity affect bee and wasp communities and their natural enemies across forest strata? Forest Ecology and Management 258: 609–615.

Vilà M, Bartomeus I, Dietzsch AC, Petanidou T, Steffan-Dewenter I, Stout JC, Tscheulin T (2009) Invasive plant integration into native plant-pollinator networks across Europe. Proceedings of the Royal Society B 276: 3887-3893.

Westphal C, Steffan-Dewenter I, Tscharntke T (2009) Mass flowering oilseed rape improves early colony growth but not sexual reproduction of bumblebees. Journal of Applied Ecology 46: 187-193. 


Bos MM, Tylianakis JM, Steffan-Dewenter I, Tscharntke T (2008) The invasive Yellow Crazy Ant and the decline of forest ant diversity in Indonesian cacao agroforests. Biological Invasions 10: 1399-1409.

Höhn P, Tscharntke T, Tylianakis JM, Steffan-Dewenter I (2008) Functional group diversity of bee pollinators increases crop yield. Proceedings of the Royal Society of London B 275: 2283-2291.

Holzschuh A, Steffan-Dewenter I, Tscharntke T (2008) Agricultural landscapes with organic crops support higher pollinator diversity. Oikos 117: 354-361.

Klein A-M, Cunningham SA, Bos MM, Steffan-Dewenter I (2008) Advances in pollination ecology from tropical plantation crops. Ecology 89: 935-943.

Ricketts TH, Regetz J, Steffan-Dewenter I, Cunningham SA, Kremen C, Bogdanski A, Gemmill-Herren B, Greenleaf SS, Klein A-M, Mayfield MM, Morandin LA, Ochieng A, Potts SG, Viana BF (2008) Landscape effects on crop pollination services: are there general patterns? Ecology Letters 11: 499-515.

Ricketts TH, Regetz J, Steffan-Dewenter I, Cunningham SA, Kremen C, Bogdanski A, Gemmill-Herren B, Greenleaf SS, Klein A-M, Mayfield MM, Morandin LA, Ochieng A, Potts SG, Viana BF (2008)  Landscape effects on crop pollination services: are there general patterns? (vol 11, pg 499, 2008); Corrigendum. Ecology Letters 11: 1121.

Steffan-Dewenter I, Schiele S (2008) Do resources or natural enemies drive bee population dynamics in fragmented habitats? Ecology 89: 1375-1387.

Steffan-Dewenter I, Westphal C (2008) The interplay of pollinator diversity, pollination services and landscape change. Journal of Applied Ecology 45: 737-741.

Thies C, Steffan-Dewenter I, Tscharntke (2008) Agriculture, Interannual landscape changes influence plant–herbivore–parasitoid interactions. Agriculture, Ecosystems and Environment 125: 266–268.

Westphal C, Bommarco R, Carré G, Lamborn E, Morison N, Petanidou T, Potts SG, Roberts SPM, Szentgyörgyi H, Tscheulin T, Vaissiére BE, Woyciechowski M, Biesmeijer JC, Kunin WE, Settele J, Steffan-Dewenter I (2008) Measuring bee diversity in different European habitats and biogeographical regions. Ecological Monographs 78: 653-671.


Bos MM, Höhn P, Saleh S, Büche B, Buchori D, Steffan-Dewenter I, Tscharntke T (2007) Insect diversity responses to forest conversion and agroforestry management. In: Tscharntke T, Leuschner C, Zeller M, Guhardja E, Bidin A (eds) The stability of tropical rainforest margins, linking ecological, economic and social constraints of land use and conservation. Springer, Berlin, 279-296.

Bos MM, Steffan-Dewenter I, Tscharntke T (2007) The contribution of cacao agroforests to the conservation of lower canopy ant and beetle diversity in Indonesia. Biodiversity and Conservation 16: 2429-2444.

Bos MM, Steffan-Dewenter I, Tscharntke T (2007) Shade tree management affects fruit abortion, insect pests and pathogens of cacao. Agriculture, Ecosystems & Environment 120: 201-205.

Bos MM, Veddeler D, Bogdanski AK, Klein A-M, Tscharntke T, Steffan-Dewenter I, Tylianakis JM (2007) Caveats to quantifying ecosystem services: Fruit abortion blurs benefits from crop pollination. Ecological Applications 17: 1841-1849.

Clough Y, Holzschuh A, Gabriel D, Purtauf T, Kruess A, Steffan-Dewenter I, Kleijn D, Tscharntke T (2007) Alpha and beta diversity of arthropods and plants in organically and conventionally managed wheat fields. Journal of Applied Ecology 44: 804-812.

Hambäck PA, Summerville KS, Steffan-Dewenter I, Krauss J, Englund G, Crist TO (2007) Habitat specialization, body size, and family identity explain lepidopteran density-area relationships in a cross-continental comparison. PNAS 104: 8368-8373.

Herrmann F, Westphal C, Moritz RFA, Steffan-Dewenter I (2007) Genetic diversity and mass resources promote colony size and forager densities of a social bee (Bombus pascuorum) in agricultural landscapes. Molecular Ecology 16: 1167-1178.

Holzschuh A, Steffan-Dewenter I, Kleijn D, Tscharntke T (2007) Diversity of flower-visiting bees in cereal fields: effects of farming system, landscape composition and regional context. Journal of Applied Ecology 44: 41-49.

Klein A-M, Vaissière BE, Cane JH, Steffan-Dewenter I, Cunningham SA, Kremen C, Tscharntke T (2007) Importance of pollinators in changing landscapes for world crops. Proceedings of the Royal Society B 274, 303-313.

Kremen C, Williams N, Aizen MA, Gemmill-Herren B, LeBuhn G, Minckley R, Packer L, Potts S, Roulston T, Steffan-Dewenter I, Vazquez D, Winfree R, Adams L, Crone E, Greenleaf S, Keitt T, Klein A-M, Regetz J, Ricketts T (2007) Pollination and other ecosystem services produced by mobile organisms: a conceptual framework for the effects of land-use change. Ecology Letters 10: 299-314.

Meyer B, Gaebele V, Steffan-Dewenter I (2007) Patch size and landscape effects on pollinators and seed set of the Horseshoe Vetch, Hippocrepis comosa, in an agricultural landscape of Central Europe. Entomologia Generalis 30: 173-185.

Meyer B, Kröger J, Steffan-Dewenter I (2007) Contribution of diurnal and nocturnal pollinators to the reproductive success of the orchid species Gymnadenia conopsea. Entomologia Generalis 30: 299-300.

Poveda K, Steffan-Dewenter I, Scheu S, Tscharntke T (2007) Plant-mediated interactions between below- and aboveground processes: decomposition, herbivory, parasitism and pollination. In: Ohgushi T, Craig T, Price P (eds) Ecological Communities: Plant Mediation in Indirect Interaction Webs. Cambridge University Press, Cambridge, 147-163.

Priess JA, Mimler M, Klein A-M, Schwarze S, Tscharntke T, Steffan-Dewenter I (2007) Linking deforestation scenarios to pollination services and economic returns in coffee agroforestry systems. Ecological Applications 17: 407-417.

Steffan-Dewenter I Kessler M, Barkmann J, Bos MM, Buchori D, Erasmi S, Faust H, Gerold G, Glenk K, Gradstein SR, Guhardja E, Harteveld M, Hertel D, Höhn P, Kappas M, Köhler S, Leuschner C, Maertens M, Marggraf R, Migge-Kleian S, Mogea J, Pitopang R, Schaefer M, Schwarze S, Sporn SG, Steingrebe A, Tjitrosoedirdjo SS, Tjitrosoemito S, Twele A, Weber R, Woltmann L, Zeller M, Tscharntke T (2007) Tradeoffs between income, biodiversity, and ecosystem functioning during tropical rainforest conversion and agroforestry intensification. PNAS 104: 4973-4978.


Kleijn D, Baquero RA, Clough Y, Diaz M, De Esteban J, Fernandez F, Gabriel D, Herzog F, Holzschuh A, Johl R, Knop E, Kruess A, Marshall EJP, Steffan-Dewenter I, Tscharntke T, Verhulst J, West TM, Yela JL (2006) A rejoinder to the comments by Potts et al.. Ecology Letters 9: 256-257.

Kleijn D, Baquero RA, Clough Y, Diaz M, De Esteban J, Fernandez F, Gabriel D, Herzog F, Holzschuh A, Johl R, Knop E, Kruess A, Marshall EJP, Steffan-Dewenter I, Tscharntke T, Verhulst J, West TM, Yela JL (2006) Mixed biodiversity benefits of agri-environment schemes in five European countries. Ecology Letters 9: 243-254. (highlighted in Nature (2006) 439, 908-909).

Klein A-M, Steffan-Dewenter I, Tscharntke T (2006) Rain forest promotes trophic interactions and diversity of trap-nesting Hymenoptera in adjacent agroforestry. Journal of Animal Ecology 75: 315-323. 

Poveda K, Steffan-Dewenter I, Scheu S, Tscharntke T (2006) Belowground effects of organic and conventional farming on aboveground plant-herbivore and plant-pathogen interactions. Agriculture, Ecosystems & Environment 113: 162-167.

Steffan-Dewenter I, Klein A-M, Gaebele V, Alfert T, Tscharntke T (2006) Bee diversity and plant-pollinator interactions in fragmented landscapes. In: Waser NM, Ollerton J (eds). Plant-pollinator interactions: from specialization to generalization. University of Chicago Press, Chicago, 387-407.

Westphal C, Steffan-Dewenter I, Tscharntke T (2006) Foraging trip duration of bumblebees in relation to landscape-wide resource availability. Ecological Entomology 31: 389-394.

Westphal C, Steffan-Dewenter I, Tscharntke T (2006) Bumblebees experience landscapes at different spatial scales: possible implications for coexistence. Oecologia 149: 289-300.


Steffan-Dewenter I, Potts SG, Packer L (2005) Pollinator diversity and crop pollination services are at risk. TRENDS in Ecology and Evolution 20: 651-652.

Krauss J, Steffan-Dewenter I, Müller CB, Tscharntke T (2005) Relative importance of resource quantity, isolation and habitat quality for landscape distribution of a monophagous butterfly. Ecography 28: 465-474.

Poveda K, Steffan-Dewenter I, Scheu S, Tscharntke T (2005) Floral trait expression and plant fitness in response to below- and aboveground plant- animal interactions. Perspectives in Plant Ecology, Evolution and Systematics 7: 77-83.

Poveda K, Steffan-Dewenter I, Scheu S, Tscharntke T (2005) Effects of decomposers and herbivores on plant performance and aboveground plant-insect interactions. Oikos 108: 503-510.

Settele J, Hammen V, Hulme P, Karlson U, Klotz S, Kotarac M, Kunin WE, Marion G, O'Connor M, Petanidou T, Peterson K, Potts S, Pritchard H, Pyšek P, Rounsevell M, Spangenberg J, Steffan-Dewenter I, Sykes M, Vighi M, Zobel M, Kühn I (2005) ALARM: Assessing LArge-scale environmental Risks for biodiversity with tested Methods - The concept, objectives, structure and management of a large Integrated Project within the 6th framework programme of the European Commission. Gaia 14: 69-72.

Tscharntke T, Klein A-M, Kruess A, Steffan-Dewenter I, Thies C (2005) Landscape perspectives on agricultural intensification and biodiversity - ecosystem service management. Ecology Letters 8: 857-874.

Veddeler D, Schulze CH, Steffan-Dewenter I, Buchori D, Tscharntke T (2005) The contribution of tropical secondary forest fragments to the conservation of fruit-feeding butterflies: effects of isolation and age. Biodiversity and Conservation 14: 3577-3592.


Klein A-M, Steffan-Dewenter I, Tscharntke T (2004) Foraging trip duration and density of megachilid bees, eumenid wasps and pompilid wasps in tropical agroforestry systems. Journal of Animal Ecology 73: 517-525.

Krauss, J, Klein A-M, Steffan-Dewenter I, Tscharntke T (2004) Effects of habitat area, isolation, and landscape diversity on plant species richness of calcareous grasslands. Biodiversity and Conservation 13: 1427-1439.

Krauss J, Schmitt T, Seitz A, Steffan-Dewenter I, Tscharntke T (2004) Effects of habitat fragmentation on the genetic structure of the monophagous butterfly Polyommatus coridon along its northern range margin. Molecular Ecology 13: 311-320.

Krauss J, Steffan-Dewenter I, Tscharntke T (2004) Landscape occupancy and local population size depends on host plant distribution in the butterfly Cupido minimus. Biological Conservation 120: 355-361.

Schulze CH, Steffan-Dewenter I, Tscharntke T (2004) Effects of land use on butterfly communities at the rain forest margin: a case study from Central Sulawesi. In: Land Use, Nature Conservation and the Stability of Rainforest Margins in Southeast Asia. Gerold G, Fremerey M, Guhardja E (eds). Springer-Verlag, Berlin, Heidelberg, New York, 281-297.

Schulze CH, Waltert M, Kessler PJA, Pitopang R, Shahabuddin, Veddeler D, Mühlenberg M, Gradstein SR, Leuschner C, Steffan-Dewenter I, Tscharntke T (2004) Biodiversity indicator groups of tropical land-use systems: comparing plants, birds, and insects. Ecological Applications 14: 1321-1333.

Steffan-Dewenter I, Schiele S (2004) Nest site fidelity, body weight and population size of the red mason bee Osmia rufa (Hymenoptera: Megachilidae), evaluated by mark-recapture experiments. Entomologia Generalis 27: 123-131. 


Klein A-M, Steffan-Dewenter I, Tscharntke T (2003) Fruit set of highland coffee increases with the diversity of pollinating bees. Proceedings of the Royal Society of London B 270: 955-961.

Klein A-M, Steffan-Dewenter I, Tscharntke T (2003) Bee pollination and fruit set of Coffea arabica and C. canephora (Rubiaceae). American Journal of Botany 90: 153-157.

Klein A-M, Steffan-Dewenter I, Tscharntke T (2003) Pollination of Coffea canephora in relation to local and regional agroforestry management. Journal of Applied Ecology 40: 837-845.

Krauss J, Steffan-Dewenter I, Tscharntke T (2003) How does landscape context contribute to effects of habitat fragmentation on diversity and population density of butterflies? Journal of Biogeography 30: 889-900

Krauss, J, Steffan-Dewenter I, Tscharntke T (2003) Local species immigration, extinction, and turnover of butterflies in relation to habitat area and habitat isolation. Oecologia 137: 591-602.

Poveda K, Steffan-Dewenter I, Scheu S, Tscharntke T (2003) Effects of below- and above-ground herbivores on plant growth, flower visitation and seed set. Oecologia 135: 601-605.

Steffan-Dewenter I (2003) Seed set of male-sterile and male-fertile oilseed rape (Brassica napus) in relation to pollinator density. Apidologie 34: 227-235.

Steffan-Dewenter I (2003) Artenvielfalt und Pflanze-Insekt-Interaktionen auf Kalkmagerrasen. Bauhinia 17: 64-65.

Steffan-Dewenter I (2003) Importance of habitat area and landscape context for species richness of bees and wasps in fragmented orchard meadows. Conservation Biology 17: 1036-1044.

Steffan-Dewenter I, Kuhn A (2003) Honeybee foraging in differentially structured landscapes. Proceedings of the Royal Society B 270: 569-575.

Steffan-Dewenter I, Leschke K (2003) Effects of habitat management on vegetation and above-ground nesting bees and wasps of orchard meadows in Central Europe. Biodiversity and Conservation 12: 1953-1968.

Thies C, Steffan-Dewenter I, Tscharntke T (2003) Effects of landscape context on herbivory and parasitism at different spatial scales. Oikos 101: 18-25.

Tscharntke T, Klein A-M, Kruess A, Steffan-Dewenter I, Thies C, (2003) Biodiversität und Pflanze-Insekt-Interaktionen in Kulturlandschaften. In: Gradstein SR, Willmann R, Zizka G (eds). Biodiversitätsforschung. Die Entschlüsselung der Artenvielfalt in Raum und Zeit. E. Schweitzerbart'sche Verlagsbuchhandlung, Stuttgart, 171-183.

Tscharntke T, Steffan-Dewenter I, Kruess A, Thies C (2003) Grassland habitats. In: Encyclopedia of insects. Resh VC, Carde R (eds). Academic Press, San Diego, 481-485.

Westphal C, Steffan-Dewenter I, Tscharntke T (2003) Mass flowering crops enhance pollinator densities at a landscape scale. Ecology Letters 6: 961-965.


Klein A-M, Steffan-Dewenter I, Buchori D, Tscharntke T (2002) Effects of land-use intensity in tropical agroforestry systems on coffee flower-visiting and trap-nesting bees and wasps. Conservation Biology 16: 1003-1014.

Steffan-Dewenter I (2002) Landscape context affects trap-nesting bees, wasps, and their natural enemies. Ecological Entomology 27: 631-637.

Steffan-Dewenter I, Münzenberg U, Bürger C, Thies C, Tscharntke T (2002) Scale-dependent effects of landscape context on three pollinator guilds. Ecology 83: 1421-1432.

Steffan-Dewenter I, Tscharntke T (2002) Insect communities and biotic interactions on fragmented calcareous grasslands - a mini review. Biological Conservation 104: 275-284.

Tscharntke T, Bürger C, Eber S, Gabriel D, Grabe H, Grönmeier M, Holzschuh A, Jahn S, Klein A-M, Klipfel S, Kluth S, Krauss J, Kruess A, Lehmann K, Poveda K, Roschewitz I, Schiele S, Schmidt M, Schulze CH, Steffan-Dewenter I, Sundmacher F, Thies C, Veddeler D, Westphal C (2002) Pflanze-Insekt-Interaktionen und Biodiversität in der Agrarlandschaft, Forschungsmagazin Georgia Augusta: Vielfalt des Lebens - Biodiversität, 1, 41-46.

Tscharntke T, Steffan-Dewenter I, Kruess A, Thies C (2002) Characteristics of insect populations on habitat fragments: a mini review. Ecological Research 17: 229-239.

Tscharntke T, Steffan-Dewenter I, Kruess A, Thies C (2002) Contribution of small habitat fragments to conservation of insect communities of grassland-cropland landscapes. Ecological Applications 12: 354-363.


Alfert T, Steffan-Dewenter I, Tscharntke T (2001) Bienen und Wespen (Hymenoptera, Aculeata) in Kalksteinbrüchen: Der Effekt von Flächengröße und Flächenalter (Bees and wasps (Hymenoptera, Aculeata) in limestone quarries: The effects of area and age). Mitteilungen der Deutschen Gesellschaft für allgemeine und angewandte Entomologie 13: 579-582.

Klein A-M, Steffan-Dewenter I, Tscharntke T (2001) Predator-prey ratios on cocoa along a land-use gradient in Indonesia. Biodiversity and Conservation 11: 683-693.

Krauss J, Steffan-Dewenter I, Tscharntke T (2001) Tagfalter auf fragmentierten Kalkmagerrasen - Hat sich die Lebensgemeinschaftsstruktur in den letzten Jahren geändert? (Butterflies on fragmented calcareous grasslands - Are there changes in community structure during the last years?). Mitteilungen der Deutschen Gesellschaft für allgemeine und angewandte Entomologie 13: 525-528.

Steffan-Dewenter I, Münzenberg U, Tscharntke T (2001) Pollination, seed set and seed predation on a landscape scale. Proceedings of the Royal Society of London B 268: 1685-1690.

Steffan-Dewenter I, Tscharntke T (2001): Succession of bee communities on fallows. Ecography 24: 83-93.


Steffan-Dewenter I, Tscharntke, T (2000) Butterfly community structure in fragmented habitats. Ecology Letters 3: 449-456.

Steffan-Dewenter I, Tscharntke T (2000) Resource overlap and possible competition between honey bees and wild bees in central Europe. Oecologia 122: 288-296.


Steffan-Dewenter I, Tscharntke T (1999) Effects of habitat isolation on pollinator communities and seed set. Oecologia 121: 432-440.


Tscharntke T, Gathmann A, Steffan-Dewenter I (1998) Bioindication using trap-nesting bees and wasps and their natural enemies: community structure and interactions. Journal of Applied Ecology 35: 708-719.


Steffan-Dewenter I, Tscharntke T (1997) Early succession of butterfly and plant communities on set-aside fields. Oecologia 109: 294-302.

Steffan-Dewenter I, Tscharntke T (1997) Bee diversity and seed set in fragmented habitats. Acta Horticulturae 437: 231-234.


Steffan-Dewenter I, Tscharntke T (1996) Profitieren Wildbienen oder Honigbienen von der Flächenstillegung in der Landwirtschaft? Natur und Landschaft 71: 255-261.

Tscharntke T, Greiler H-J, Steffan-Dewenter I, Kruess A, Gathmann A, Zabel J, Wesserling J, Dubbert M, Kuhnhenne J, Vu M-H (1996) Die Flächenstillegung in der Landwirtschaft - eine Chance für Flora und Fauna der Agrarlandschaft? (Set-aside in agriculture - a chance for plants and animals of the agricultural landscape?) NNA-Berichte 2/96: 59-72.


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